Academic Education

• 1964          A.B. (cum laude), University of Pennsylvania, The College


• 1968          M.D., University of Pennsylvania, School of Medicine, Philadelphia, PA


• 1968-69     Internship in Medicine, University of California, San Francisco, CA


• 1972-74     Residency in Neurology, University of California, San Francisco, CA

Appointments and Professional Activities

• 1974-80     Assistant Professor of Neurology in Residence, Univ of California, SF


• 1976-88     Lecturer, Department of Biochemistry and Biophysics, UCSF


• 1979-83     Assistant Professor of Virology in Residence, UC, Berkeley


• 1980-81     Associate Professor of Neurology in Residence, UCSF


• 1981-84     Associate Professor of Neurology, UCSF


• 1983-84     Associate Professor of Virology in Residence, UC, Berkeley


• 1984-         Professor of Neurology, UCSF


• 1984-         Professor of Virology in Residence, UC, Berkeley


• 1988-         Professor of Biochemistry, UCSF

Major Honors and Awards

• Potamkin Prize for Alzheimer's Disease Research, American Academy of Neurology, 1991


• Christopher Columbus Quincentennial Discovery Award in Biomedical Research, NIH, 1992 Metropolitan Life Foundation Award for Medical Research, 1992


• Dickson Prize for Distinguished Scientific Accomplishments, University of Pittsburgh, 1992


• Charles A. Dana Award for Pioneering Achievements in Health, 1992


• Richard Lounsbery Award for Extraordinary Scientific Research, NAS, 1993


• Gairdner Foundation Award for Outstanding Achievement in Medical Science, 1993


• Bristol-Myers Squibb Award for Distinguished Achievement in Neuroscience Res., 1994


• Albert Lasker Award for Basic Medical Research, Albert and Mary Lasker Foundation, 1994


• Paul Ehrlich Prize, Paul Ehrlich Foundation and the Federal Republic of Germany, 1995


• Wolf Prize in Medicine, Wolf Foundation and the State of Israel, 1996


• Keio International Award for Medical Science, Keio University, Tokyo, Japan, 1996